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Published papers

  1. May, K.A. & Zhaoping, L. (2016). Efficient coding theory predicts a tilt aftereffect from viewing untilted patterns. Current Biology, 26, 1571–1576.
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  2. Solomon, J.A., May, K.A. & Tyler, C.W. (2016). Inefficiency of orientation averaging: Evidence for hybrid serial/parallel temporal integration. Journal of Vision, 16(1):13, 1–7.
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  3. May, K.A. & Solomon, J.A. (2015). Connecting psychophysical performance to neuronal response properties II: Contrast decoding and detection. Journal of Vision, 15(6):9, 1–21.
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  4. May, K.A. & Solomon, J.A. (2015). Connecting psychophysical performance to neuronal response properties I: Discrimination of suprathreshold stimuli. Journal of Vision, 15(6):8, 1–26.
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  5. Hansen,B.C., May, K.A. & Hess, R.F. (2014). One "shape" fits all: The orientation bandwidth of contour integration. Journal of Vision, 14(13):17, 1–21.
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  6. Dumoulin, S.O., Hess, R.F., May, K.A., Harvey, B.M, Roker, B. & Barendregt, M. (2014). Contour extracting networks in early extrastriate cortex. Journal of Vision, 14(5):18, 1–14.
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  7. May, K.A. & Solomon, J.A. (2013). Four theorems on the psychometric function. PLoS ONE, 8(10):e74815.
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  8. May, K.A. & Zhaoping, L. (2013). The potential roles of saturating and supersaturating contrast-response functions in conjunction detection: Reply to Peirce. Journal of Vision, 13(4):22, 1–4.

  9. McIlhagga, W.H. & May, K.A. (2012). Optimal edge filters explain human blur detection. Journal of Vision, 12(10):9, 1–13.
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  10. Huang, P.-C., Maehara,G., May, K.A. & Hess, R.F. (2012). Pattern masking: The importance of remote spatial frequencies and their phase alignment. Journal of Vision, 12(2):14, 1–13.
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  11. May, K.A., Zhaoping, L. & Hibbard, P.B. (2012). Perceived direction of motion determined by adaptation to static binocular images. Current Biology, 22, 28–32.  Download PDF (588 KB)  Supplemental Experimental Procedures (56 KB)
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  12. May, K.A. & Zhaoping, L. (2011). Exploring the roles of saturating and supersaturating contrast-response functions in conjunction detection and contrast coding. Journal of Vision, 11(9):11, 1–15.
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  13. Zhaoping, L., Geisler, W.S. & May, K.A. (2011). Human wavelength discrimination of monochromatic light explained by optimal wavelength decoding of light of unknown intensity. PLoS ONE, 6(5): e19248.
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  14. May, K.A. & Zhaoping, L. (2009). Effects of surrounding frame on visual search for vertical or tilted bars. Journal of Vision, 9(13):20, 1–19.
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  15. May, K.A. & Hess, R.F. (2008). Effects of element separation and carrier wavelength on detection of snakes and ladders: Implications for models of contour integration. Journal of Vision, 8(13):4, 1–23.  Download Quicktime Movie (4.5 MB)
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  16. Hess, R.F., Baker, D.H., May, K.A. & Wang, J. (2008). On the decline of 1st and 2nd order sensitivity with eccentricity. Journal of Vision, 8(1):19, 1–12.
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  17. May, K.A. & Hess, R.F. (2007). Ladder contours are undetectable in the periphery: A crowding effect? Journal of Vision, 7(13):9, 1–15.
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  18. May, K.A. & Hess, R.F. (2007). Dynamics of snakes and ladders. Journal of Vision, 7(12):13, 1–9.
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  19. Georgeson, M.A., May, K.A., Freeman, T.C.A. & Hesse, G.S. (2007). From filters to features: Scale–space analysis of edge and blur coding in human vision. Journal of Vision, 7(13):7, 1–21.
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  20. May, K.A. & Georgeson, M.A. (2007). Added luminance ramp alters perceived edge blur and contrast: A critical test for derivative-based models of edge coding. Vision Research, 47(13), 1721–1731.  Download PDF (305 KB)  Supplementary files (5 KB)
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  21. May, K.A. & Georgeson, M.A. (2007). Blurred edges look faint, and faint edges look sharp: The effect of a gradient threshold in a multi-scale edge coding model. Vision Research, 47(13), 1705–1720.  Download PDF (352 KB)  Supplementary files (4 KB)
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  22. Zhaoping, L. & May, K.A. (2007). Psychophysical tests of the hypothesis of a bottom-up saliency map in primary visual cortex. PLoS Computational Biology, 3(4), e62.  Download PDF (2.3 MB) Note: This pdf file has higher-quality images than the one that is now available on the PLoS Computational Biology website. They used to provide high and low quality versions, but now only provide low-quality versions.
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  23. Perrett, D.I., May, K.A. & Yoshikawa, S. (1994). Facial shape and judgements of female attractiveness. Nature, 368, 239–242.  Download PDF (501 KB)  Nature cover (111 KB)
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  24. Thompson, P., May, K. & Stone, R. (1993). Chromostereopsis: A multicomponent depth effect? Displays, 14, 227–234.  Download PDF (1.4 MB)
    This paper is also avaliable in CBZ format, in both high quality (10.8 MB) and medium quality (3.6 MB). Note: CBZ files are actually zip files containing the scanned images. You can change the file extension to '.zip' and unzip them as normal to get the image files. But if you keep them as CBZ files, you can read them with the free CDisplay comic book reader, which is a lot less sluggish than Acrobat Reader when viewing scanned documents.
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Published book chapters

  1. Hess, R.F., May, K.A. & Dumoulin, S.O. (2015). Contour Integration: Psychophysical, neurophysiological and computational perspectives. In Wagemans, J. (Ed), The Oxford Handbook of Perceptual Organization. Oxford: Oxford University Press.  Download PDF (693 KB)

  2. Zhaoping, L., May, K.A. & Koene, A. (2009). Some fingerprints of V1 mechanisms in the bottom up saliency for visual selection. In Heinke, D. & Mavritsaki, E. (Eds.), Computational Modelling in Behavioural Neuroscience: Closing the gap between neurophysiology and behaviour (pp. 137–164). London: Psychology Press. ISBN 978-1-84169-738-3.  Download PDF (301 KB)

  3. Zhaoping, L. & May, K.A. (2008). Testing the hypothesis that V1 creates a bottom-up saliency map. In Bharath, A. & Petrou, M. (Eds.), Next Generation Artificial Vision Systems: Reverse Engineering the Human Visual System. London: Artech House. ISBN 978-1-59693-224-1.  Download PDF (547 KB)

Published conference abstracts


  1. May, K.A. & Georgeson, M.A. (in press). An optimal estimator for edge contrast explains perceived contrast of sine wave gratings. Perception. (Poster presented at AVA Christmas Meeting 2016)  Download poster (1.48 MB)
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  2. May, K.A. & Zhaoping, L. (in press). Adaptation of face gender, expression, and head direction from random-noise adaptation images: A surprising prediction of Li and Atick's efficient binocular coding theory. Perception, 45(S1), ECVP Abstract Supplement. (Talk presented at ECVP 2016)
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  3. May, K.A. & Zhaoping, L. (2016). Face gender adaptation from random noise adaptors: A surprising prediction of Li and Atick's efficient binocular coding theory. Journal of Vision, 16(12):1326. (Talk presented at VSS 2016)
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  4. May, K.A. & Zhaoping, L. (in press). Tilt aftereffect generated by isotropic adaptation stimuli: A counterintuitive prediction of efficient coding theory. i-Perception (Talk presented at SVG 2015)
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  5. May, K.A. & Zhaoping, L. (2016). Perceived direction of tilt determined by adaptation to unoriented or untilted binocular stimuli: Surprising predictions of efficient coding theory. Perception, 45, 694. (Talk presented at AVA Christmas Meeting 2015)
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  6. May, K.A. & Zhaoping, L. (2015). Tilt aftereffect generated by isotropic adaptation stimuli: A counterintuitive prediction of Li and Atick's efficient binocular coding theory. Perception, 44(S1), ECVP Abstract Supplement, 184. (Talk presented at ECVP 2015)
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  7. May, K.A. & Solomon, J.A. (2014). Formal relationships between neuronal response properties and psychophysical performance. i-Perception, 5(5):56. (Talk presented at SVG 2014)
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  8. May, K.A. & Solomon, J.A. (2013). Reconciling multiplicative physiological noise and additive psychophysical noise. Perception, 42, ECVP Abstract Supplement, 158. (Talk presented at ECVP 2013 symposium, Visual Noise: New Insights)
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  9. May, K.A. & Solomon, J.A. (2012). Weibull β for contrast detection is the Naka-Rushton exponent. Perception, 41, 1523. (Talk presented at AVA Christmas meeting 2012)
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  10. May, K.A. & Zhaoping, L. (2011). Contrast-response functions, Fisher information, and contrast decoding performance. Journal of Vision, 11(11):1172. (Poster presented at VSS 2011)  Download poster (1.67 MB)
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  11. Zeiner, K.M., Spitschan, M., May, K.A., Zhaoping, L. & Harris, J.M. (2011). Eye movements and reaction times for detecting monocular regions in binocularly viewed scenes. Journal of Vision, 11(11):326. (Poster presented at VSS 2011)
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  12. May, K.A. (2011). Bayesian decoding of neural population responses explains many characteristics of contrast detection and discrimination. Perception, 40, ECVP Abstract Supplement, 51. (Talk presented at ECVP 2011)
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  13. Dumoulin, S.O., Hess, R.F., Harvey, B.M. & May, K.A. (2010). Measuring contour integration mechanisms using fMRI. Society for Neuroscience Abstracts, 531.9.
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  14. May, K.A., Zhaoping, L. & Hibbard, P.B. (2010). Binocular integration in human vision adapts to maximize information coding efficiency. Perception, 39, ECVP Abstract Supplement, 77. (Poster presented at ECVP 2010)  Download poster (16.9 MB)
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  15. May, K.A., Zhaoping, L. & Hibbard, P.B. (2010). Binocular integration in human vision adapts quickly to maximize coding efficiency. Perception, 39, 1149. (Talk presented at the AVA AGM 2010)
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  16. Zhaoping, L. & May, K.A. (2010). Human monochromatic light discrimination explained by optimal signal decoding. Perception, 39, 1148–1149. (Talk presented at the AVA AGM 2010)
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  17. May, K.A., Zhaoping, L. & Hibbard, P.B. (2010). Effects of image statistics on stereo coding in human vision. Journal of Vision, 10(7):359. (Poster presented at VSS 2010)  Download poster (5.3 MB)
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  18. May, K.A. & Hess, R.F. (2010). Implementing curve detectors for contour integration. Perception, 39(2), 270. (Poster presented at the AVA Christmas meeting 2009)  Download poster (15.7 MB)
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  19. May, K.A. & Hess, R.F. (2009). Implementing curve detectors for contour integration. Journal of Vision, 9(8):906, 906a. (Poster presented at VSS 2009)  Download poster (15.7 MB)
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  20. May, K.A. & McIlhagga, W.H. (2009). Probing edge blur perception with reverse correlation. Perception, 38(4), 621. (Talk presented at the AVA AGM 2009)
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  21. May, K.A. & Hess, R.F. (2008). Testing filter-overlap models of contour integration. Journal of Vision, 8(6):72, 72a. (Talk presented at VSS 2008)
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  22. May, K.A. & Hess, R.F. (2007). Contour integration and crowding: a similar type of mechanism? Perception, 36(9), 1399. (Talk presented at the AVA AGM 2007)
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  23. May, K.A. & Hess, R.F. (2007). Ladder contours are undetectable in the periphery. Journal of Vision, 7(9):113, 113a. (Talk presented at VSS 2007)
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  24. May, K.A. & Hess, R.F. (2006). Snakes are as fast as ladders: evidence against the hypothesis that contrast facilitation mediates contour detection. Journal of Vision, 6(6), 337a. (Poster presented at VSS 2006) Download Poster as  Powerpoint file (4.1 MB) or  A3 size pdf (4.4 MB)
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  25. May, K.A. & Zhaoping, L. (2005). Both cognitive factors and local inhibition mediate the effect of a surrounding frame in visual search for oriented bars. Journal of Vision, 5(8), 959a. (Poster presented at VSS 2005)  Download poster (819 KB)
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  26. Guyader, N., May, K.A. & Zhaoping, L. (2005). Top-down interference in visual search. Journal of Vision, 5(8), 951a. (Poster presented at VSS 2005)
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  27. Zhaoping, L. & May, K. (2004). Irrelevance of feature maps for bottom up visual saliency in segmentation and search tasks. Society for Neuroscience Abstracts, 20.1.
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  28. May, K.A. & Zhaoping, L. (2004). Investigating salience mechanisms by using the effects of surrounding frame on the tilted-vertical asymmetry in visual search. Perception, 33, Supplement, 12. (Talk presented at ECVP 2004)
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  29. May, K.A. & Georgeson, M.A. (2004). Perceiving edge contrast. Perception, 33(6), 757. (Talk presented at the AVA Christmas Meeting 2003)
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  30. May, K.A. & Georgeson, M.A. (2003). Perceiving edge blur: Gaussian-derivative filtering and a rectifying nonlinearity. Perception, 32, Supplement, 46. (Talk presented at ECVP 2003)
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  31. May, K.A. & Georgeson, M.A. (2003). Perceiving edge blur: linear filtering and a rectifying nonlinearity. Perception, 32(3), 388. (Talk presented at the AVA Christmas Meeting 2002)
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  32. Georgeson, M.A., May, K.A. & Barbieri-Hesse, G.S. (2003). Perceiving edge blur: the Gaussian-derivative template model. Journal of Vision, 3(9), 360a.
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Conference presentations without published abstracts

  1. May, K.A. & Zhaoping, L. (2015). Tilt aftereffect from isotropic adaptators and motion aftereffect from static adaptors: Counterintuitive predictions of Li and Atick's efficient binocular coding theory. (Poster presented at Second ViiHM Workshop, July 2015, Bath, UK)  Download poster (3.36 MB)
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  2. May, K.A. (2015). Predicting neuronal decoding performance using Fisher information (Talk presented at ViiHM Mini-workshop, January 2015, City University London).

  3. May, K.A. (2014). Understanding visual coding: Insights from psychophysics and computational theory. (Keynote talk given at the 2014 Applied Vision Association Christmas meeting on receipt of the David Marr Medal)
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  4. May, K.A. (2014). Edge detection and contour integration in human and machine vision. (Poster presented at First ViiHM Workshop, September 2014, Stratford-Upon-Avon, UK)  Download poster (1.53 MB)
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  5. May, K.A., Zhaoping, L. & Hibbard, P.B. (2012). Motion adaptation from static binocular images: A surprising prediction of efficient coding theory. (Poster presented at Sensory Coding & Naural Environment 2012, IST Austria)  Download poster (12.8 MB)
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  6. May,K.A. (2010). Effects of image statistics on stereo coding in human vision. (Talk presented at the Scottish Vision Group meeting, March 2010)

  7. May, K.A. & Perrett,D.I. (1993). Facial Attractiveness. (Talk presented at the BPS International Conference on Face Processing, University of Wales College of Cardiff, September 1993)
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  8. May, K.A. (1993). Face representation. (Talk presented at SERC Workshop on Spatial Representation, Pitlochry, March 1993)


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